English: Cabbage order.
Content: glucosinolate !!, myrosin; myrosinase, erucic acid, eicosenoic acid.
Subphases1. Capparidaceae,
Cleomaceae,
Resedaceae,
Gyrostemonaceae,
Tovariaceae, Pentadiplandraceae,
Akaniaceae,
Tropaeolaceae, Limnanthaceae
2. Aetionemoideae
3.
Heliophylloideae4.
Brassicoideae5.
Cardaminoideae6. Hesperoideae
7.
Caricaceae, Moringaceae,
Salvadoraceae, Bataceae, Koeberlinaceae
TaxonomyThe order
Brassicales consist of 17 families.
Brassicaceae is the best known family and contains the majority of species of this order. In older classifications the main families Capparcaeae and
Brassicaceae were placed together in Capparales.
In the Apg2 established the order. Assembled DNA sequence data sets for glucosinolate taxa and applied cladistic methods showed that all the glucosinolate-producing taxa form a clade, the
Brassicales, save Drypetes. Despite the paucity of morphological synapomorphies the chemistry and DNA show a well supported evidence for the group’s monophyly.
In older classifications Limnanthaceae had been placed in its own order, Limnanthales.
Tropaeolaceae in
Geraniales, Batidaceae in
Caryophyllales, Caricacaeae in
Violales.
Plant TheoryIn the first version of the Plan theory The main
Family Brassicaceae was placed in
Subphase 6. Many smaller Families occupied the other
Subphases.
In this update
Brassicaceae is split in 5 Subfamilies, which are placed in
Subphase 2 to
Subphase 6. In
Subphase 1 is placed
Capparaceae with many other small families. It makes
Subphase 1 not monophyletic. Capparidaceae,
Cleomaceae have long be considered as one family, but have been split due to reasons of monophyly.
Resedaceae,
Gyrostemonaceae,
Tovariaceae and Pentadiplandraceae form a monophyletic clade.
Akaniaceae and
Tropaeolaceae form a monophyletic clade. Limnanthaceae is a family on itds own with one species.
Caricaceae is the main
Family in
Subphase 7.
Caricaceae and Moringaceae for a monophyletic clade and
Salvadoraceae, Bataceae, Koeberlinaceae form another monophyletic clade. Together they miss the monophyly. But they both have
Phase 7 qualities.
Families:
Akaniaceae, Bataceae,
Caricaceae (
Carica papaya), Emblingiaceae,
Gyrostemonaceae,
Koeberliniaceae, Limnanthaceae, Moringaceae (
Moringa oleifera), Pentadiplandraceae,
Resedaceae,
Salvadoraceae, Setchellantacaee,
Tovariaceae,
Tropaeolaceae (
Tropaeolum majus).
LiteratureBailey, C. Donovan & others; Toward a Global
Phylogeny of the
Brassicaceae.
Beilstein, Mark A & others; BRASSICACEAE PHYLOGENY INFERRED FROM PHYTOCHROME A AND NDHF SEQUENCE DATA: TRIBES AND TRICHOMES REVISITED; American Journal of
Botany 95,10: 1307–1327; 2008.
Franzke, Andreas & others; Arabidopsis family ties: molecular phylogeny and age estimates in
Brassicaceae; Taxon 58, 2: 425–437; May 2009.
Franzke, Andreas & others; Cabbage family affairs: the evolutionary history of
Brassicaceae; Trends in plant science, Volume 16,2: p108–116, February 2011.
Andreas FranzkeEmail the author Andreas Franzke, Martin A. Lysak, Ihsan A. Al-Shehbaz, Marcus A. Koch, Klaus Mummenhoff
Hall, Jocelyn & others; PHYLOGENY OF CAPPARACEAE AND BRASSICACEAE BASED ON CHLOROPLAST SEQUENCE DATA; American Journal of
Botany 89,11: 1826–1842; 2002.
Hall, Jocelyn & others; Molecular phylogenetics of core
Brassicales; Systemcic
Botany vol 29.3: 654-669; 2004.
Kagale, Sateesh & othres; Polyploid Evolution of the
Brassicaceae during the Cenozoic Era; The Plant Cell, Vol. 26: 2777–2791; July 2014.