7 Pteridophyta TaxonomyIntroductionEnglish: Ferns; Pteridophytes.
Synonym: Pteridopsida;
Cryptogamae.
Content: thiaminase; tannins; flavonoids.
Members: Ferns, Horsetails, Lycophytes, Clubmosses, Spikemosses, Quillworts.
Plant theoryPteridophyta is a phylum in the Plant kingdom, including the Ferns, Horsetails and Clubmosses. It is the first clade in the
Tracheophyta, the vascular plants.
Pteridophyta are placed in the Phylum 4, having the connection with the
Iron series.
The classification of the ferns has been difficult. Firstly, there was hardly any information in homeopathy about them.
Lycopodium clavatum was the exception, but is was so overused and has such a vast
Materia medica that it is hard to find the essence in it.
ClassificationThere are four classes in the
Pteridophyta and ten subclasses. The 10 subclasses have each one order, there is only
Phase 1.
This expresses that all ferns have an impulsive quality. They just do things, without thinking about it, only following the rules. It is a kind of naivety. It is also an expression of their being young in evolution. They are more children and teenager remedies, even when much needed in adults, for their child and teenager personalities. It is as if there is no past, only a future.
The opposite is the case for
Fungi, where there are 7 orders, each in one of the 7
Phases in a subclass, but each order has only 1 family and thus 1 Subphase. This expresses that
Fungi has much past, but little future.
Table Subclasses & OrdersNr
Series Remedy code
Classes Subclasses Order
411 Hydrogen 3-411.10.00
Isoetanae Isoetidae Isoetales421 Carbon 3-421.10.00
Lycopodianae Salviniidae Salviniales422 Carbon 3-422.10.00
Lycopodianae Lycopodiidae Lycopodiales431 Silicon 3-431.10.00
Equisetanae Azollidae Azollales432 Silicon 3-432.10.00
Equisetanae Equisetidae Equisetales433 Silicon 3-433.10.00
Equisetanae Cyatheidae Cyatheales441 Iron 3-441.10.00
Polypodianae Marsileidae Marsileales442 Iron 3-442.10.10
Polypodianae Pterididae Pteridales443 Iron 3-443.10.00
Polypodianae Aspleniidae Aspleniales444 Iron 3-444.10.00
Polypodianae Polypodiidae PolypodialesTaxonomyPteridophyta are not monophyletic because the Monilophyta, ferns and horsetails, are more closely related to seed plants than to Lycophyta.
Pteridophyta is thus not widely accepted as a taxon anymore, but remains in common parlance, as do pteridology and pteridologist. This is so as
Pteridophyta share a life cycle and are collectively treated or studied, for example by the International Association of Pteridologists and the Pteridophyte
Phylogeny Group. A nice example is the
Pteridophyta itself: it is not monophyletic as the
Spermatophyta are arising in them, in between the clubmosses and the real fern, the Euphyllophyta. But the PPG group itself is treating them both. This problem arises from using the criterion of monophyletic too strictly.
The botanical classification of ferns was not very developed. This changed very much with the publication on ferns by the Pteridophyte
Phylogeny Group, PPG 1, in 2016. This made the homeopathic classification much easier. The table below shows the relationships between the orders as in the
Plant theory with the families of the PPG1 classification.
Nr Order Families411
Isoetales Isoetaceae.
421
Salviniales Salviniaceae.
422
Lycopodiales Lycopodiaceae,
Huperziaceae,
Selaginellaceae.
431
Azollales Azollaceae.
432
Equisetales Psilotaceae,
Ophioglossaceae,
Equisetaceae, Marratiaceae,
Osmundacae,
Schizaeaceae,
Lygodiaceae.
433
Cyatheales Hymenophyllaceae,
Gleicheniaceae,
Matoniaceae,
Dipteridaceae,
Cyatheaceae,
Cibotiaceae,
Culcitaceae,
Dicksoniaceae,
Anemiaceae.
441
Marsileales Marsileaceae.
442
Pteridales Lindsaeceae +,
Pteridaceae,
Dennstaedtiaceae.
443
Aspleniales Blechnaceae,
Onocleaceae,
Woodsiaceae,
Athyriaceae,
Thelypteridaceae, Cystopterridaceae +,
Aspleniaceae.
444
Polypodiales Hypodematiaeae,
Didymochlaenaceae,
Dryopteridaceae,
Polypodiaceae,
Tectariaceae,
Davalliaceae,
Oleandraceae,
Lomariopsidaceae,
Nephrolepidaceae.
Discussion differencesIn general the botanical and the
Plant theory classification are very similar.
Phase 3 families with their theme of confusion.
Fusion of cladesSome clades are fused in the
Plant theory, which is in contradiction with the monophyletic in the PPG classification:
Equisetidae,
Cyatheidae,
Pterididae.
Some Families have absorbed smaller neighbours, without losing their monophyly:
Gleicheniaceae,
Culcitaceae,
Dicksoniaceae,
Lindsaeaceae,
Thelypteridaceae,
Dryopteridaceae and
Polypodiaceae.
In several cases the "outsider"
Phases,
Phase 1 and
Phase 7, got included in the main clade. For instance,
Lindsaeaceae and
Dennstaedtiaceae got included in
Pteridales. As outsiders they are more different in their character, morphology and DNA than the central families.
Splitting of clades
In some cases clades have been split, mostly in such a way that the secludes are also monophyletic:
Lycopodiaceae,
Huperziaceae,
Pteridaceae,
Dryopteridaceae,
Polypodiaceae.
The biggest change is the split of
Salviniales in 3 quite separate subclasses.
Differences Plant theory and PPG1classificationNr Order Families
411
Isoetales Isoetaceae, without
Selaginellaceae.
421
Salviniales Salviniaceae, shifted to
Lycopodianae,
without
Azollaceae and
Marsileaceae.
422
Lycopodiales Lycopodiaceae split in 3,
Huperziaceae split in 3.
with
Selaginellaceae.
431
Azollales Azollaceae separated from
Salviniales.
432
Equisetales Paraphyletic due to the fusion of the families.
Ophioglossaceae split in 2.
Schizaeaceae without
Anemiaceae.
433
Cyatheales Paraphyletic due to the fusion of the families.
Gleicheniaceae including
Matoniaceae,
Dipteridaceae,
Culcitaceae +
Plagiogyriaceae,
Loxsomataceae,
Thyrsopteridaceae.
Dicksoniaceae including Metaxyaceae.
Anemiaceae split off from
Schizaeales.
441
Marsileales Marsileaceae separated from
Salviniales.
442
Pteridales Paraphyletic, including
Lindsaeaceae,
Dennstaedtiaceae.
Lindsaeceae including Cystodiaceae,
Lonchitidaceae,
Saccolomataceae.
Pteridaceae split in 5.
443
Aspleniales Aspleniidae = PPG1 Aspleniineae, = Eupolypods 2.
Onocleaceae including
Woodsiaceae.
Thelypteridaceae split in 2.
Cystopterridaceae + Desmophlebiaceae, Dplaziopsidaceae,
Hemidictyaceae,
Rhachidosoraceae.
444
Polypodiales Polypodiidae = PPG1 Polypodiineae, =Eupolypods 1.
Elaphoglossoideae + Hypodematiaeae,
Didymochlaenaceae.
Dryopteridaceae, split in 2
Polypodiaceae, split in 4.
Tectariaceae +
Davalliaceae,
Oleandraceae,
Lomariopsidaceae,
NephrolepidaceaeNomenclatureIn the
Plant theory somewhat different names are given to classes and subclasses. In the PPG classification classes and subclasses are not exactly recognised, as is done in the
Plant theory. In accordance with the endings of clades classes are given the ending "-anae" and subclasses "-idae". In the PPG classification subclasses are ending in "ineae".
Classes are difficult to recognise and if so get the ending "-opsida". This last ending can be confusing as some families have a similar ending, like Berberidopsis.
The inconsistency is obvious for derivations of Polypodium. Polypodiopsida is a kind of superclass for all
Pteridophyta without the clubmosses.
Polypodiidae is a kind of superclass for all Polypodiopsida without the
Equisetidae. The biggest subgroup of Polypodiopsida is
Polypodiales, which has the ending of an order, but contains 3 subclasses. One of them is the subclass Polypodiineae.
BotanyFerns, Horsetails and Clubmosses have vessels for the first time in the evolution of plants. They are vascular plants together with later evolutionary clades: the
Spermatophyta. Hence the name
Tracheophyta, vessel plants. The vessels make transport possible and with that specialisation in the form of roots, leaves, stems and reproduction systems. It can make them bigger than for instance mosses. This aspect of transport and specialisation is an aspect of the
Iron series.
Ferns have having xylem and phloem, making them more rigid and woody, and thus making them bigger.
The reproduction is via spores. As spore plants they have no flowers or seeds.