PortulacalesSynonym: Portulacaceous alliance; Portulacinae; Portulacaceous cohort.
Clades:
Caryophyllidae,
Asteranae,
Angiospermae,
Plants.
BotanyHerbs; many succulents.
TaxonomyPortulacales is a clade in the
Apg4 classification, a Suborder in the Order
Caryophyllales. It is called the Potulacoid clade. It contains the Families:
Anacampserotaceae,
Basellaceae Cactaceae, Didieraceae,
Halophytaceae,
Montiaceae,
Portulacaceae and
Talinaceae.
Cactaceae is by far the biggest family in
Portulacales. In good right the Order could be called Cactales. Recently there have been made big shift in some families. The quite big family
Portulacaceae lost most of its members to
Montiaceae and has only Portulaca as its member.
Plant theoryIn the
Plant theory Portulacales is an Order in the Subclass of
Caryophyllidae and is placed in
Phase 1. The placement in Sub- phases is tentative. In the update of the
Plant theory The Portulalcales is reordered. Some small families have been fused.
In
Subphase 1 is combined Didieraceae and
Halophytaceae.
In
Subphase 2 is combined
Portulacaceae,
Anacampserotaceae,
Talinaceae and
Basellaceae.
In
Subphase 3 is
Montiaceae.
The
Cactaceae is split in its 4 Subfamilies:
Echinocactoideae,
Phyllocactoideae,
Opuntioideae and Cereoioideae respectively in
Subphase 4 to 7.
Subphases1.
Halophytaceae, Didiereacaeae
2.
Portulacaceae3.
Montiaceae4.
Echinocactoideae,
Cactaceae5.
Phyllocactoideae,
Cactaceae6.
Opuntioideae,
Cactaceae7. Cereoioideae,
CactaceaeSubphases1.
Halophytaceae, Didiereacaeae
2.
Portulacaceae, including
Anacampserotaceae,
Basellaceae,
Talinaceae,
Didiereaceae: they feel alone in relationships, adapt too much to the other.
3.
Montiaceae: they stay single, in triangular relationships.
4. Echoinocactoideae: they prefer to stay single as others are imperfect.
5.
Phyllocactoideae.
6.
Opuntioideae: they prefer to stay single from fear of being used.
7.
Cereoideae.
LiteratureNyffeler, Reto & Eggli, Urs; A farewell to dated ideas and concepts – molecular phylogenetics and a revised suprageneric classification of the family
Cactaceae; Zurich Open Repository and Archive; 2010.