Author: Jan Scholten
The Caryophyllaceae were not well divided in Subphases in the first version of the Plant theory. The lack of information and remedies made that too speculative. Recent information and rpovings of new remedies has made the division possible.
The placement of Caryophyllaceae in Phase 4 has been quite well established. The care for their family is a keynote for the Caryophyllaceae and they do not question that.
Caryophyllaceae is traditionally divided in three subfamilies:
- Alsinoideae: no stipules, petals not united.
Silenoideae or Caryophyllideae: no stipules, petals united.
- tribe Caryophylleae.
- tribe Sileneae
Paronychioideae: fleshy stipules, petals separate or united.
- tribe Paronycheae
- tribe Polycarpeae
The tribe Caryophylleae with Dianthus, Gypsophyla and Saponaria have the most stable form in their family situation. Their vision of the family is sure, indicating Subphase 4.
Sileneae, the other tribe in Silenoideae, shows aspects of Subphase 6. Cases show absent parents and parents neglecting their children.
The next tribe becoming clear was Paronycheae in Subphase 7. Cases of Herniaria and Paroncyhia showed suicide of the father, a beating father, a mother with insanity and suicidal tendency or being an unwanted child.
Paronychioideae are a basal grade of rather primitive members Caryophyllaceae, not closely related with many plesiomorphic traits. This indicates Subphases 1 and 7, at the start or end. They are not central. This is another confirmation for Paronychieae in Subphase 7.
Polycarpea eis the other tribe Paronychioideae, even more plesiomorphic with Corrigiola as a first line in it. This can indicate Subphase 1. This si confirmed with a case of Spergularia and the marine quality fs some Spergularia plants.
The subfamily Alsinoideae is not easily divided in tribes. The biggest Genus is Arenaria, but that has to be split in some different Genera according to Fior.
Proving of Cerastium and Eregenome have indications for Subphase 3. Stellaria cases also have a Subphase 3 qualities. Another case of Stellaria shows has foster children, a theme of Subphase 3. The clade in Fior containing those genera, Group B, is taken as Subphase 3. I have given it the name Stellarieae.
Scleranthus is known for his timidity and Calcium like aspects, Subphase 2. A proving of Minuartia confirmed this. This clade, In Fior without a name, is given the name Sclerantheae, the name of a subtribe.
The last clade, Group C in Fior, is then left for Subphase 5. There is though a lack of information to confirm this.
1. Polycarpeae: Spergularia, Corrigiola, Drymaria
2. Sclerantheae: Scleranthus, Minuartia, Drypis. ( Bufonia, Sagina )
3. Stellarieae: Stellaria, Cerastium, Eregenome, Holosteum.
4. Caryophylleae: Dianthus, Gypsophyla, Saponaria.
5. Alsineae: Arenaria, Moehringia.
6. Sileneae: SIlene, Lychnis, Agrostema.
7. Paronychieae: Paronychia, Herniaria.
Fior, Simone & Karis, Per Ola & Casazza, Gabriele & Minuto, Luigi & Sala, Fransesco; MOLECULAR PHYLOGENY OF THE CARYOPHYLLACEAE (CARYOPHYLLALES) INFERRED FROM CHLOROPLAST MATK AND NUCLEAR RDNA ITS SEQUENCES; American Journal of Botany 93(3); 399'96411; 2006.